Document Type

Article

Publication Date

1-1-2011

Publication Title

Genetica

Abstract

The multiple species concepts currently in use by the scientific community (e. g. Morphological, Biological, Phylogenetic) are united in that they all aim to capture the process of divergence between populations. For example, the Biological Species Concept defines a species as a natural group of organisms that is reproductively isolated from other such groups. Here we synthesize nearly a century of research on the ciliate genus Paramecium that highlights the shortcomings of our prevailing notions on the nature of species. In this lineage, there is discordance between morphology, mating behavior, and genetics, features assumed to be correlated, at least after sufficient time has passed, under all species concepts. Intriguingly, epigenetic phenomena are well documented in ciliates where they influence features such as germline/soma differentiation and mating type determination. Consequently, we hypothesize that divergence within ciliate populations is due to a dynamic interaction between genetic and epigenetic factors. The growing list of examples of epigenetic phenomena that potentially impact speciation (i. e. by influencing the dynamics of sex chromosomes, fate of hybrids, zygotic drive and genomic conflicts) suggests that interactions between genetics and epigenetics may also drive divergence in other eukaryotic lineages.

Keywords

Ciliates, Epigenetics, Molecular evolution, Paramecium, Species concepts

Volume

139

Issue

5

First Page

677

Last Page

684

DOI

10.1007/s10709-011-9571-3

ISSN

00166707

Rights

© 2011 Springer Science+Business Media B.V.

Comments

Peer reviewed accepted manuscript.

Included in

Biology Commons

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