Document Type

Article

Publication Date

4-1-2008

Publication Title

Molecular Biology and Evolution

Abstract

Analyses of diverse eukaryotes reveal that genomes are dynamic, sometimes dramatically so. In numerous lineages across the eukaryotic tree of life, DNA content varies within individuals throughout life cycles and among individuals within species. Discovery of examples of genome dynamism is accelerating as genome sequences are completed from diverse eukaryotes. Though much is known about genomes in animals, fungi, and plants, these lineages represent only 3 of the 60-200 lineages of eukaryotes. Here, we discuss diverse genomic strategies in exemplar eukaryotic lineages, including numerous microbial eukaryotes, to reveal dramatic variation that challenges established views of genome evolution. For example, in the life cycle of some members of the "radiolaria," ploidy increases from haploid (N) to approximately 1,000N, whereas intrapopulation variability of the enteric parasite Entamoeba ranges from 4N to 40N. Variation has also been found within our own species, with substantial differences in both gene content and chromosome lengths between individuals. Data on the dynamic nature of genomes shift the perception of the genome from being fixed and characteristic of a species (typological) to plastic due to variation within and between species.

Keywords

Epigenetics, Genome evolution, Genome rearrangements, Life cycle variation, Microbial eukaryotes, Polyploidy

Volume

25

Issue

4

First Page

787

Last Page

794

DOI

10.1093/molbev/msn032

ISSN

07374038

Rights

© The Author 2008.

Comments

Peer reviewed accepted manuscript.

Included in

Biology Commons

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